한국에서는 제주도형이 오랫동안 L. makinoana의 학명을 사용했으며, 이는 내륙에 나타나는 두 종 중 하나인 L. longiracemosa로도 인식되었다. 그러나 내륙형 중 하나는 후에 원래의 L. makinoana로 재확인되었다. 형태학적 특성으로 볼 때, 이 종은 L. longiracemosa와 L. suzumushi 사이의 특성을 가지고 있다. 이 식물은 비교적 크기가 크며, 10-35cm이다. 잎은 타원형이거나 타원형-타원형이며, 잎 가장자리는 통일되어 있다. 꽃은 완만하게 피어나며, 꽃대에 5-35개가 핀다. 입술은 반원형이며, 8-12 × 5-9mm 크기이며, 다른 두 종과 비교하여 초록색, 황록색 또는 보라색을 띤다. Liparis makinoana와 L. longiracemosa는 6월부터 7월까지 꽃을 피우지만, L. makinoana는 L. longiracemosa의 꽃 피는 시기보다 2~3주 늦게 피어난다. L. makinoana는 해발 60-900m 지역에 자생지가 있다. 가장 낮은 해발 지역은 충청남도의 연안 산맥으로, 가장 높은 지역은 강원도의 산림지역이다. 이 종은 주로 한국의 중북부 지역에 분포하며, 분포 지역을 포함하여 L. longiracemosa와 구별된다.
Description
Terrestrial herbs, 10–35 cm tall. Roots ca. 1 mm in diam. Pseudobulbs aggregated, ovoid, 1–3 × 0.5–3.5 cm, enclosed by 2 or 3 membranaceous sheaths; sheaths ovate-lanceolate, 1–3 cm long, shortly subacute at apex. Leaves 2; petiole base sheathing, enclosing peduncle, winged, 3–9 cm long; blades green, elliptic or ovate-elliptic, slightly obtuse, 10–20 × 2.5–8 cm, conduplicate, plicate, apex slightly obtuse, margin entire, occasionally undulate, both surfaces glabrous. Inflorescence terminal, racemose, 10–30 cm long; scape slightly slender, relatively long, glabrous, ca. 5–20 cm long, ca. 2.5–4.5 mm in diam., slightly compressed cylindric, narrow or broad wings on either side, 3(4)-ridged, green; laxly bearing 5–35 flowers; floral bracts elliptic to ovate, triangular, 1.0–5.5 mm long, subacute to acute, green. Pedicel and ovary twisted at base, ca. 10–18 mm long, green, greenish-purple, or purplish. Flowers yellowish-green, green, purple, or dark-purple, widely open, green or purple especially in middle of lip; dorsal sepal linear-lanceolate, 9–13 × 2–3 mm, apex subacute, slightly revolute, erect or sometimes recurved, yellowish-green to purplish; lateral sepals yellowish-green to purple, obliquely elliptic or oblong-lanceolate, 9–13 × 2–3 mm, apex subacute, sometimes revolute; petals greenish or purplish, falcate, 8–12 × 0.5–1.0 mm, linear, obtuse, apex slightly revolute, pendulous, sometimes slightly twisted, positioned lower than lateral sepals; lip porrect, obovate, 8–12 × 5–9 mm, narrowed toward base, apex roundly truncate and mucronate, minutely apiculate in the middle of the apex, clawed, strongly recurved near base, green, yellowish-green, or purple. Column green to purple-tinged, incurved, with rounded wings at apex, 3.5–5.0 mm long, much dilated at base, pale-green to green on ventral surface, green to purple in the basal part; anther cap ovate, 2-celled, cells surrounded by whitish rim, with beaked in front, ligule-like caudate, slightly curved, obliquely rounded, become narrow and slightly obtuse at the end part, green; pollinia 4 in 2 pairs, ovoid, deltoid, waxy, yellow. Capsules ellipsoid, 2.5–3.5 × 0.5–0.7 cm, stalk 3–8 mm long
Distribution
Korea. Gangwon-do: Inje, Hongcheon, Pyeongchang, Jeongseon, Samcheok. Gyeonggi-do: Pocheon. Chungcheongnam-do: Taean. Yanggang-do: Mt. Baekdu Yalu-river. Japan. Hokkaido, Honshu. China. Jilin, Heilongjiang. Russia. Far East
Note
Liparis makinoana was designated as a neotype based on the protologue, morphology, and molecular studies of the original description, and the prototype of L. makinoana was amended. An herbarium sheet (LE 01016040) collected near Hakodate, Hokkaido, Japan, was designated as a neotype of L. makinoana. In Fig. 8, the plant on the left was designated as a type plant of L. makinoana (Tsutsumi et al., 2019), according to Art. 9.17 of the current International Code of Nomenclature for algae, fungi, and plants (Turland et al., 2018). The large plant in the center is L. koreojaponica Tsutsumi, T. Yukawa, N. S. Lee, C. S. Lee et M. Kato, and the small plants on the periphery it is thought to be L. makinoana or L. suzumushi (Tsutsumi et al., 2019), but a detailed examination of these plants is still required. In Korea, the Jeju type has long been assigned the scientific name of L. makinoana, and it was also recognized as L. longiracemosa, one of two species appearing inland. However, one of the inland types was subsequently reidentified as the original L. makinoana (Figs. 7–9). In terms of its morphological traits, this species has traits between those of L. longiracemosa and L. suzumushi. These plants are relatively tall at 10–35 cm. Leaves are elliptic or ovate-elliptic; the leaf margin is entire. The flowers are laxly borne, with 5–35 flowered on the rachis. The lip is obovate, 8–12 × 5–9 mm, and has a green, yellowish-green, or purple color compared to the other two species. Liparis makinoana and L. longiracemosa have a similar flowering time of June–July, but L. makinoana has a flowering time two to three weeks later than that of L. longiracemosa. L. makinoana is native to altitudes of 60–900 m. The area with the lowest altitude consisted of coastal hills in Chungcheongnam-do, and the highest area was a mountain forest in Gangwon-do. This species is mainly distributed in the mid-northern region of Korea and is distinct from L. longiracemosa, including in its distribution area. As with L. longiracemosa and L. suzumushi, we propose that the familiar “Na-li-nan-cho” should continue to be used. “Na-li-nan-cho” is the original national name for L. makinoana.
Reference
Tsutsumi, C., T. Yukawa and M. Kato. 2019. Taxonomic reappraisal of Liparis japonica and L. makinoana (Orchidaceae). Bulletin of National Museum of Nature and Science, Series B, Botany 45: 107–118.
Lee, J. and D.C.Son. 2023. Taxonomic revision of the Liparis makinoana complex (Orchidaceae; Epidendroideae; Malaxidae) in Korea. Korean Journal of Plant Taxonomy 53(2): 110-125.
Vegetative character
Liparis makinoana grows in semi-open deciduous and mixed forests, forest edges, glades.
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